Physiology and Ethology of Domestic Animals

A.Y. 2025/2026
12
Max ECTS
128
Overall hours
SSD
VET/02 VET/08
Language
Italian
Learning objectives
The course is divided into three modules. The first module, Physiology of nervous, cardiocirculatory and muscular system (6 CFU), provides students with a comprehensive overview of cellular function physiology, the nervous system and sensory organs, the blood and cardiovascular system, and the skeletal and smooth muscle cells in domestic mammals, laboratory animals, fish, and birds. Upon completion of the module, students will be able to apply the knowledge acquired to understand the key physiological mechanisms underlying the functioning of organs and the systems involved. They will also be able to analyze and address physiological issues relevant to veterinary clinical practice. The second module, Veterinary ethology and animal welfare (4 CFU), provides students with foundational knowledge and skills related to the behavior of domestic animals, including specific references to certain species of new companion animals (NAC). Upon completion of the module, students will be able to identify and correctly interpret the legal provisions relevant to the veterinary profession, including regulations on animal protection and welfare, as well as offenses committed against animals. Additionally, they will develop skills useful for managing situations that require the proper application of veterinary law in a professional context. The third module, Veterinary Law (2 CFU), aims to provide students with a foundational understanding of national and European law relevant to veterinary practice. Upon completion of the module, students will be able to identify and correctly interpret the legal provisions relevant to the veterinary profession, including regulations on animal protection and welfare, as well as offenses committed against animals. Additionally, they will develop skills useful for managing situations that require the proper application of veterinary law in a professional context.
Expected learning outcomes
1) Knowledge and understanding: for the physiology module, students should be able to explain the basic physiological functions related to the nervous system, including neuron physiology, the spinal cord, brain (particularly the cerebellum and motor functions), and sensory organs. They should also be able to describe the basic functions of the cardiovascular system, including cardiac physiology (electrocardiography and mechanical cardiac activity with nervous control) and circulatory physiology (relationship between pressure, flow, and resistance across all circulatory districts and nervous regulation of pressure), highlighting differences among domestic mammals, birds, and fish. Additionally, students should understand the physiological bases of skeletal, smooth, and cardiac muscle contraction in domestic mammals, fish, and birds. For the ethology module: the student is expected to demonstrate knowledge of the physiological foundations of animal behavior, with a particular focus on underlying mechanisms, development, communication, and cognitive processes. Additionally, the student must demonstrate the ability to scientifically study animal behavior. For the law module, students should acquire a basic understanding of law, the veterinarian's relationship with the judicial authority and other competent bodies and institutions, knowledge of animal welfare laws, and regulations on animal protection and welfare.
2) Applying knowledge and understanding: students must demonstrate the ability to connect various topics, such as the functions of the nervous system, particularly the hypothalamus, and different animal behaviors, such as feeding behavior or responses to temperature changes. This will enable them to adequately address subsequent studies in pathology and veterinary clinic. Students should apply ethological knowledge to improve the welfare of domestic animals, including NAC. They must also develop the ability to critically search and analyze relevant legislation and case law and refer to the judicial authority, preparing them for further study and understanding of veterinary legal medicine and legislation.
3) Critical thinking and judgment: the student will be expected to approach the study of the subject matter critically. This will include demonstrating the ability to orally present written reports during practical activities, integrating the study materials provided by the professor of physiology with independently acquired knowledge. Additionally, the student will need to present the ethological studies conducted in the veterinary ethology module. The student will also be expected to critically engage with the study of law, identifying relevant regulations pertaining to professional practice in various fields of interest, evaluating and correctly applying them, and recognizing the associated responsibilities.
4) Communication: students will need to demonstrate the use of appropriate terminology to adequately express acquired knowledge. This will be shown through responses to open questions during exams, class presentations of reports prepared during the course (which will be evaluated by professors), and active participation in class discussions, including arguing cases proposed by the professor.
5) Life-long learning skills: students will be expected to demonstrate their ability to independently expand their knowledge of topics related to physiology, ethology, and the welfare of domestic species, including exotic companion animals (NAC), as well as veterinary law.

This ability will be assessed through practical activities where students will be assigned small research projects requiring them to independently consult up-to-date scientific and legal sources (such as articles, guidelines, codes, and regulations) to address a given problem or explore a specific topic in depth. Assessment will focus on the quality of the sources selected, the student's ability to analyze the content, and their capacity to apply the information gathered to develop relevant solutions or interpretations.
Single course

This course cannot be attended as a single course. Please check our list of single courses to find the ones available for enrolment.

Course syllabus and organization

Single session

Responsible
Lesson period
First semester
Prerequisites for admission
The prerequisites required to pass the exam are the knowledge of the anatomy of the nervous, cardiocirculatory and skeletal muscular systems, as well as the basics of biochemistry. It is also necessary to have passed the preliminary examinations of:
1) Biochemistry and Molecular Biology and 2) Systematic and Comparative Veterinary Anatomy.
Assessment methods and Criteria
The exam consists of three written tests: one for physiology, one for ethology, and one for veterinary law.
The physiology test is divided into two sections: 1) 20 multiple-choice questions each with 4 answer options. Each correct option is worth 0.25 points, up to a maximum of 1 point per question. Questions may have more than one correct answer. 2) 2 open-ended questions. Each can receive a score from 0 to 5 points, based on the completeness of the answer (0-2 points), the accuracy of the content (0-2 points), and the appropriate use of scientific language (1 point). The final exam score will be the sum of the points obtained in both sections. The ethology test consists of 15 multiple-choice questions, each with four possible answers, only one of which is correct. Each correct answer is worth 2 points, while incorrect or unanswered questions do not receive any points. The maximum score for this test is 30 points.
The Veterinary Law test consists of 8 multiple-choice questions, each with four possible answers, only one of which is correct; two of the questions are followed by an open-ended in-depth question. Each correct answer is worth 3.25 points. Each answer to the in-depth questions is worth between 0 and 2 points, depending on the accuracy and clarity of the answer. Wrong answers or no answers are not penalised. The maximum score for this test is 30 points. All three tests are conducted at the same time. The final exam grade is calculated as a weighted average of the scores obtained in each module. Additionally, up to one extra point may be awarded based on the evaluation of presentations submitted during practical exercises.
Students will have a total of three hours to complete the exam.
Phisiology of the nervous, cardiovascular and muscular system
Course syllabus
CELL PHYSIOLOGY: plasmatic membrane and membrane transports: 2 hours
CENTRAL AND PERIPHERAL NERVOUS SYSTEM PHYSIOLOGY in domestic species and laboratory animals.
Neurons: resting and action membrane potential, impulse conduction: 2 hours
Synapses and neurotransmitters: 1 hour
Receptor classification; tactile, pain and temperature receptors: 2 hours
Spinal cord structure and function: spinal reflexes: 1 hour
Cranial nerve functions: 1 hour
Brainstem: functions and autonomic centers: 1 hour
Muscle tone control: 1 hour
Labirintic reflexes: 1 hour
Cerebellum: 1 hour
Thalamus and hypothalamus: 2 hours
Cortical areas of the brain and EEG: 1 hour
Autonomic nervous system: 2 hours
Sensory organs in domestic mammals, fish and birds: gustatory, olfactory, auditory and vision: 3 hours
MUSCLE ACTIVITY in domestic mammals, fish and birds.
Skeletal muscle: structure, excitation-contraction coupling: 2 hours
Neuromuscular transmission: 1 hour
Smooth muscle: 1 hour
Muscle tissue in birds and fish: 1 hour
BLOOD
Blood function and composition: 1 hour
Red blood cell parameters: 1 hour
Haematopoiesis and haemostasis: 1 hour
CARDIOCIRCULATORY PHYSIOLOGY in domestic mammals, fish and birds.
Physiology of cardiac cells: conduction tissue and working myocardium: 1 hour
Electrocardiography: cardiac electrical axis and ECG: 1 hour
Cardiac cycle, cardiac output and heart sounds: 2 hours
Nervous control of cardiac function: 1 hour
Basics of circulatory mechanics: 1 hour
Venous and arterial systems; arterial pressure: 1 hour
Microcirculation and capillary pressure: Starling's hypothesis: 1 hour
Central and peripheral control of circulation: 1 hour.
Cardiovascular function in fish: 1 hour.
Cardiovascular function in birds: 1 hour.

1) Classroom exercises:
2 hours for 4 groups of 25 students each: explanation of how to conduct a bibliographic search using the main available databases and how to properly construct a scientific paper. Selection of a scientific paper to be critically analyzed by subgroups of up to 3 students each. Subsequently, 9 hours with all students present: presentation and analysis of the scientific paper selected by student subgroups, focusing on the physiology of domestic or non-conventional animals.
2) Exercises at a dog breeding facility:
4 hours for 4 groups of 25 students each: correct approach to dogs and measurement of main physiological parameters, e.g., heart rate, heart sounds, and nervous reflexes (e.g., palpebral and miotic reflexes; patellar reflex).
3) Laboratory Exercises:
1 hour for 4 groups of 25 students each: demonstration of how to correctly perform an ECG.
Teaching methods
Frontal lessons through projection of slides with diagrams, figures, text and explanatory videos. At the end of each lesson, IT tools (Wooclap) are used to verify the understanding of the topics covered, also allowing students to formulate questions anonymously.
The exercises take place partly in the classroom, partly in lab and partly in the field.
1) Classroom exercises:
2 hours to 4 groups of 25 students each.
9 hours in the presence of all the students.
2) Visits to a canine training center:
4 hours to 4 groups of 25 students each.
3) Exercises in Lab:
1 hour to 4 groups of 25 students each.
Teaching Resources
Albertini et al., Fisiologia Veterinaria. Point Veterinaire Italie, 2024.
Sjaastad, Sand e Hove, Fisiologia degli animali domestici. Casa Editrice Ambrosiana, 2013.
Slides and videos of the course are available on the Ariel website.
Veterinary ethology and animal welfare
Course syllabus
Introduction to ethology with a historical overview, role of ethology in veterinary medicine: 2 hours
Genetic-based behaviors and animal mind: 4 hours
Learning Systems: 4 hours
Ontogeny of behavior: 3 hours
Social system of the species covered in the course: 8 hours
The study of behavior: 3 hours

1) Classroom practicals
N. 5 hours conducted with the entire class present: presentation of ethological studies carried out by students working in groups of 4.
2) Field practicals
N. 4 hours for 4 groups of 25 students at a canine training center: activities focused on social behavior, personality and mental activation.
N. 3 hours in the presence of all students at the horse paddock in Lodi: observation of behavior, personality, and cognition.
3) Laboratory practicals at the Animal Physio-Ethology Lab (VET/02) in Lodi:
N. 4 hours for 4 groups of 25 students: canine olfaction and cognition.
Teaching methods
The lectures are delivered using slides that include diagrams, images, text, and explanatory videos. Digital tools such as Wooclap and Google Forms are used to assess learning progress, encourage interactivity, and gather feedback from students.
The practicals are conducted partly in the classroom, and partly in the field and laboratory.
1) Classroom practicals:
N. 5 hours in the presence of all students.
2) Field practicals:
N. 4 hours divided into 4 groups of 25 students each at a dog training center.
N. 3 hours with all students at a horse paddock or stable.
3) Laboratory practicals:
N. 4 hours divided into 4 groups of 25 students each.
Teaching Resources
M. Albertini, E. Canali, S. Cannas, V. Ferrante, S. Mattiello, M. Panzera, M. Verga, Etologia applicata e benessere animale, Vol. 1 e 2. Point Veterinaire Italie, 2008.
Albertini et al., Fisiologia Veterinaria. Point Veterinaire Italie, 2024.
G. Bogliani, C. Carere, R. Cervo, D. A. Grasso, P. Luschi, ETOLOGIA Lo studio del comportamento animale, UTET Università, 2022.
Slides and scientific literature available on Ariel.
Veterinary legislation
Course syllabus
General Part:
Introduction to Law: Concepts of the national and EU legal systems (2 hours)
The judicial system (1 hour)
The legal status of animals: status and classification (1 hour)
Elements of criminal and civil law for the veterinary profession Buying and selling animals and warranty (1-2 hours)
Animal welfare: general legal aspects (2 hours)

Exercises:
No. 4 hours with 4 groups of 25 students each: illustration of how to carry out legal and jurisprudential research, using the main databases available online; setting up work on a case study; problem-solving exercise, preceded by the presentation of the case, research and analysis of the application of the sectoral legislation, followed by discussion.

No. 12 hours in plenary: Critical examination of practical cases selected from case law in the light of the relevant regulations, with the participation of all students. Topics of interest to the veterinary profession will be identified:

- Animal mistreatment and other offences against animals
- Protection and welfare of farm animals
- Animal welfare and protection during transport
- Animal welfare and protection at slaughter
- Animal welfare for scientific and educational purposes; conscientious objection
- Protection of companion animals and urban hygiene issues
- Wildlife protection and hunting regulations
- Animal welfare in zoos and wildlife parks, circuses, and travelling shows
Teaching methods
Lectures (1 CFU) + exercises (1 CFU). If necessary, experts in the field can be invited for seminars or workshops. The exercises are carried out in the classroom: N. 4 hours for 4 groups of 25 students each and N. 12 hours for the whole class.
Teaching Resources
Slides provided by the lecturer and available on the MyAriel website
Reference regulations
Judgments of interest
Articles on the topics covered, published in national and international journals
Institutional websites
Modules or teaching units
Phisiology of the nervous, cardiovascular and muscular system
VET/02 - VETERINARY PHYSIOLOGY - University credits: 6
Practicals: 16 hours
Lessons: 40 hours
Shifts:
1 turno per un gruppo di studenti
Professor: Albertini Mariangela
2 turno per un gruppo di studenti
Professor: Albertini Mariangela
3 turno per un gruppo di studenti
Professor: Albertini Mariangela
4 turno per un gruppo di studenti
Professor: Albertini Mariangela
Turno unico per tutti gli studenti
Professor: Albertini Mariangela

Veterinary ethology and animal welfare
VET/02 - VETERINARY PHYSIOLOGY - University credits: 4
Practicals: 32 hours
Lessons: 16 hours
Professor: Pirrone Federica
Shifts:
Professor: Pirrone Federica
1 turno per un gruppo di studenti
Professor: Pirrone Federica
2 turno per un gruppo di studenti
Professor: Pirrone Federica
3 turno per un gruppo di studenti
Professor: Pirrone Federica
4 turno per un gruppo di studenti
Professor: Pirrone Federica
Turno unico per tutti gli studenti
Professor: Pirrone Federica

Veterinary legislation
VET/08 - VETERINARY CLINICAL MEDICINE - University credits: 2
Practicals: 16 hours
Lessons: 8 hours
Professor: Fossati Paola
Shifts:
Professor: Fossati Paola
1 turno per un gruppo di studenti
Professor: Fossati Paola
2 turno per un gruppo di studenti
Professor: Fossati Paola
3 turno per un gruppo di studenti
Professor: Fossati Paola
4 turno per un gruppo di studenti
Professor: Fossati Paola
Turno unico per tutti gli studenti
Professor: Fossati Paola